维基百科
異蟲綱
异虫纲(学名:Xenusia)是一類生存在古生代海洋中的已滅絕動物,是叶足动物的代表演化支,可能包括有爪动物的祖先。[2]模式属是异虫属(Xenusion)。[3]它们具有相对较大的环状圆柱形的身体,它们的叶足基部有结节。有些附肢较大,[3]这些埋藏的化石生物資訊能够做为其代表。[4]它们的口器在躯体末端或接近末端,它们曾经生活在全球各地的海洋中。[5]它们可能是一个並系群,而不是单系群。
分类
始有爪目 Archonychophora
- 始有爪目 Archonychophora Hou & Bergstrom, 1995[2]– undifferentiated appendages; each segment the same as each other.[6]
- 啰哩山虫科 Luolishaniidae Hou & Bergstrom, 1995 – three small rounded sclerites per segment – these are thorn-shaped spines.[6]
- 啰哩山虫属 Luolishania Hou & Chen 1989
- 神奇啰哩山虫属 Miraluolishania Liu & Shu 2004
- 贫腿虫科 Paucipodiidae Hou et al., 2004[7]– with no dorsal sclerites, two claws, fine annulation and few segments.
- 贫腿虫属 Paucipodia Hou et al. 2004
- 啰哩山虫科 Luolishaniidae Hou & Bergstrom, 1995 – three small rounded sclerites per segment – these are thorn-shaped spines.[6]
These taxa are only known from the Chengjiang (Cambrian Stage 2 Series 3).
原有爪目 Protonychophora
- 原有爪目 Protonychophora Hutchinson, 1930 – spiny legs, reduced tail.
- 埃谢栉蚕科 Aysheaiidae Walcott, 1911 – many claws on each leg. Anterior grasping appendages with long spines.
- 埃谢栉蚕属 Aysheaia Walcott 1911
- 异虫科 Xenusiidae Dzik & Krumbiegel, 1989 – >20 leg-bearing segments; paired, rounded sclerites on each segment; spiny legs.
- 异虫属 Xenusion Pompeckj, 1927
- 尖山叶足虫属 Jianshanopodia Liu et al.[3]– from Chengjiang; two rows of tubercles but no obvious sclerites.
- Hadranax Budd & Peel 1998 – from the Sirius Passet. Lacks obvious dorsal armature, but bears four 'nodes' per row.
- 埃谢栉蚕科 Aysheaiidae Walcott, 1911 – many claws on each leg. Anterior grasping appendages with long spines.
坚有爪目 Scleronychophora
- 坚有爪目 Scleronychophora Hou & Bergstrom, 1995 – paired sclerites; elongated head sclerites.
- 始小贝科 Eoconchariidae Hou & Shu, 1987 – ~10 leg-bearing segments, small head, sieve-like sclerites, varying in shape along body; curved claws on annulated legs; terminal mouth and anus.
- 微网虫属 Microdictyon Bengtson et al.
- Quadratapora Hao and Shu, 1987
- Fusuconcharium Hao and Shu, 1987
- 怪诞虫科 Hallucigeniidae Conway Morris, 1977 – ~10 leg-bearing segments; large sclerites covering head; sclerites on each segment are long spines.
- 心网虫科 Cardiodictyidae Hou & Bergstrom, 1995 – many segments (~23); large sclerites cover head; hexagonal sclerite on each body segment.
- 心网虫属 Cardiodictyon Hou et al. 1991 – known from the Chengjiang.[8]
- 始小贝科 Eoconchariidae Hou & Shu, 1987 – ~10 leg-bearing segments, small head, sieve-like sclerites, varying in shape along body; curved claws on annulated legs; terminal mouth and anus.
等有爪目 Paronychophora
- 等有爪目 Paronychophora Hou & Bergstrom, 1995 – short, downwards facing head; papillae on body and legs, some arranged in rows; shield-like sclerites on head; dorsal armature of spines;[9]claw-like jaws; annulated legs [Note: diagnosis modified in [6]].
Order unassigned
- Orstenotubulus Maas et al. 2007 – from the Furongian Orsten deposits – with retractable dorsal spines.[10]
- Carbotubulus Haug et al. 2012[11]– from the Mazon creek. Dorwal armature uncertain. Few segments, long limbs.
- Mureropodia[12]from the Stage 2 Murero lagerstatten, Spain.
'门' 有爪动物门
Onychophora are distinguished by their terrestrial habit, their ventral mouth; their antennae, jaws and oral papillae; they seem to be most closely related to the Paronychophora. Their first fossils are Carboniferous (Helenodora), although they may have had a cryptic earlier history.
參考文獻
- ^ Dzik, J.; Krumbiegel, G. N. The oldest 'onychophoran' Xenusion: A link connecting phyla?. Lethaia. 1989, 22 (2): 169–181. doi:10.1111/j.1502-3931.1989.tb01679.x.
- ^ 2.0 2.1 Hou, X.; Bergström, J. A. N. Cambrian lobopodians-ancestors of extant onychophorans?. Zoological Journal of the Linnean Society. 1995, 114: 3–19. doi:10.1111/j.1096-3642.1995.tb00110.x.
- ^ 3.0 3.1 3.2 Jianni Liu Degan Shu, Jian Han, Zhifei Zhang & Xingliang Zhang. A large xenusiid lobopod with complex appendages from the Lower Cambrian Chengjiang Lagerstätte (PDF). Acta Palaeontologica Polonica. 2006, 51 (2): 215–222 [2020-04-27]. (原始内容 (PDF)于2017-08-09).
- ^ Julián Monge-Nájera & Xianguang Hou; Hou. (PDF). Revista de Biología Tropical. 2002, 50 (3–4): 1133–1138. PMID 12947596. (原始内容 (PDF)存档于June 29, 2011).
- ^ 5.0 5.1 George Poinar, Jr. Fossil onychophorans from Dominican and Baltic amber: Tertiapatus dominicanus n.g., n.sp. (Tertiapatidae n.fam.) and Succinipatopsis balticus n.g., n.sp. (Succinipatopsidae n.fam.) with a proposed classification of the subphylum Onychophora. Invertebrate Biology. 2000, 119 (1): 104–109. JSTOR 3227105. doi:10.1111/j.1744-7410.2000.tb00178.x.
- ^ 6.0 6.1 6.2 Ma, X.; Hou, X.; Bergström, J. Morphology of Luolishania longicruris (Lower Cambrian, Chengjiang Lagerstätte, SW China) and the phylogenetic relationships within lobopodians. Arthropod Structure & Development. 2009, 38 (4): 271–291. doi:10.1016/j.asd.2009.03.001.
- ^ Xian-Guang Hou, Xiao-Ya Ma, Jie Zhao & Jan Bergström; Ma; Zhao; Bergström. The lobopodian Paucipodia inermis from the Lower Cambrian Chengjiang fauna, Yunnan, China. Lethaia. 2004, 37 (3): 235–244. doi:10.1080/00241160410006555.
- ^ 8.0 8.1 Whittle, R. J.; Gabbott, S. E.; Aldridge, R. J.; Theron, J. An Ordovician Lobopodian from the Soom Shale Lagerstätte, South Africa. Palaeontology. May 2009, 52 (3): 561–567. doi:10.1111/j.1475-4983.2009.00860.x.
- ^ Liu, J.; Shu, D.; Han, J.; Zhang, Z.; Zhang, X. The Lobopod Onychodictyon from the Lower Cambrian Chengjiang Lagerstätte Revisited. Acta Palaeontologica Polonica. 2008, 53 (2): 285–292. doi:10.4202/app.2008.0209 .
- ^ Maas, A.; Mayer, G.; Kristensen, R. M.; Waloszek, D. A Cambrian micro-lobopodian and the evolution of arthropod locomotion and reproduction. Chinese Science Bulletin. 2007, 52 (24): 3385–3392. doi:10.1007/s11434-007-0515-3.
- ^ Haug, J. T.; Mayer, G.; Haug, C.; Briggs, D. E. G. A Carboniferous Non-Onychophoran Lobopodian Reveals Long-Term Survival of a Cambrian Morphotype. Current Biology. 2012, 22 (18): 1673–1675. PMID 22885062. doi:10.1016/j.cub.2012.06.066 .
- ^ Gámez Vintaned, J. A.; Liñán, E.; Zhuravlev, A. A New Early Cambrian Lobopod-Bearing Animal (Murero, Spain) and the Problem of the Ecdysozoan Early Diversification. Evolutionary Biology – Concepts, Biodiversity, Macroevolution and Genome Evolution. 29 June 2011: 193–219. ISBN 978-3-642-20762-4. doi:10.1007/978-3-642-20763-1_12.