蜥虎属

蜥虎属（学名：Hemidactylus），又名 蝎虎屬，爬行纲有鳞目壁虎科的一属。

蜥虎属
原尾蜥虎（Hemidactylus bowringii）
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	爬行綱 Reptilia
目：	有鳞目 Squamata
科：	壁虎科 Gekkonidae
属：	蜥虎属 Hemidactylus Oken, 1817
異名^[1]
Aliurus Dunn & Dunn, 1940 Emydactylus Hemidact Oken, 1817 Hemidactylus Oken, 1817 Hemidactylys Leiurus Gray, 1845 Liurus Cope, 1862

特征

蜥虎属壁虎瞳孔垂直。指、趾明显扩展，末节侧扁，独立于扩展部，均具爪。攀蹼对分。雄性具有肛前孔、股孔或两者兼具。身体背部被粒鳞或间以较大的疣鳞。^[2]

下属物种

本属包括以下物种：^[3]

• Hemidactylus aaronbaueri Giri, 2008
• Hemidactylus acanthopholis Mirza & Sanap, 2014^[4]
• Hemidactylus achaemenidicus Torki, 2019
• Hemidactylus adensis Šmíd, Moravec, Kratochvíl, Nasher, Mazuch, Gvoždík & Carranza, 2015
• Hemidactylus aemulus Kumar, Srinivasulu & Srinivasulu, 2022^[5]
• Hemidactylus afarensis Šmíd, Mazuch, Nováková, Modry, Malonza, Abdirahman-Elmi, Carranza & Moravec, 2020
• Hemidactylus agrius Vanzolini, 1978
• Hemidactylus albituberculatus Trape, 2012
• Hemidactylus albivertebralis Trape & Böhme, 2012
• Hemidactylus albofasciatus Grandison & Soman, 1963
• Hemidactylus albopunctatus (Loveridge, 1947)
• Hemidactylus alfarraji Shobrak, Wilms, Joger & Carranza, 2016
• Hemidactylus alkiyumii Carranza & Arnold, 2012
• Hemidactylus almakhwah ^[6]
• Hemidactylus angulatus Hallowell, 1854
• Hemidactylus ansorgii Boulenger, 1901
• Hemidactylus aporus Boulenger, 1906
• 缅北蜥虎 Hemidactylus aquilonius Mcmahan & Zug, 2007
• Hemidactylus arnoldi Lanza, 1978
• Hemidactylus asirensis Šmíd, Shobrak, Wilms, Joger & Carranza, 2016^[7]
• Hemidactylus awashensis Šmíd, Moravec, Kratochv Il, Nasher, Mazuch, Gvoždík & Carranza, 2015
• Hemidactylus barbierii Sindaco, Razzetti & Ziliani, 2007
• Hemidactylus barbouri Loveridge, 1942
• Hemidactylus barodanus Boulenger, 1901
• Hemidactylus bavazzanoi Lanza, 1978
• Hemidactylus bayonii Bocage, 1893
• Hemidactylus benguellensis Bocage, 1893
• Hemidactylus beninensis Bauer, Tchibozo, Pauwels & Lenglet, 2006
• Hemidactylus biokoensis Wagner, Leaché & Fujita, 2014
• Hemidactylus boavistensis (Boulenger, 1906)
• Hemidactylus bouvieri (Bocourt, 1870)
• 原尾蜥虎 Hemidactylus bowringii (Gray, 1845)
• Hemidactylus brasilianus Amaral, 1935
• 密疣蜥虎 Hemidactylus brookii Gray, 1845
• Hemidactylus carivoensis Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Pinto, 2021
• Hemidactylus chikhaldaraensis Agarwal, Bauer, Giri & Khandekar, 2019^[8]
• Hemidactylus chipkali Mirza & Raju, 2017^[9]
• Hemidactylus cinganji Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Pinto, 2021
• Hemidactylus citernii Boulenger, 1912
• Hemidactylus coalescens Wagner, Leaché & Fujita, 2014
• Hemidactylus craspedotus Mocquard, 1890
• Hemidactylus curlei Parker, 1942
• Hemidactylus dawudazraqi Moravec, Kratochvíl, Amr, Jandzik, Šmíd & Gvoždík, 2011
• Hemidactylus depressus Gray, 1842
• Hemidactylus dracaenacolus Rösler & Wranik, 1999
• Hemidactylus easai Das, Pal, S, V.Palot & Narayanan, 2022
• Hemidactylus echinus O'Shaughnessy, 1875
• Hemidactylus endophis Carranza & Arnold, 2012
• Hemidactylus eniangii Wagner, Leaché & Fujita, 2014
• Hemidactylus farasani ^[6]
• Hemidactylus fasciatus Gray, 1842^[10]
• Hemidactylus faustus Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Pinto, 2021
• Hemidactylus festivus Carranza & Arnold, 2012
• Hemidactylus flavicauda Lajmi, Giri, Singh & Agarwal, 2020^[11]，也拼作Hemidactylus flavicaudus
• Hemidactylus flaviviridis Rüppell, 1835^[12]
• Hemidactylus forbesii Boulenger, 1899
• Hemidactylus foudaii Baha El Din, 2003
• Hemidactylus fragilis Calabresi, 1915
• 疣尾蜥虎 Hemidactylus frenatus Duméril & Bibron, 1836
• Hemidactylus funaiolii Lanza, 1978
• 锯尾蜥虎 Hemidactylus garnotii Duméril & Bibron, 1836
• Hemidactylus giganteus Stoliczka, 1871
• Hemidactylus gleadowi Murray, 1884
• 纤细蜥虎 Hemidactylus gracilis Blanford, 1870
• Hemidactylus gramineus Ceríaco, Bauer, Kusamba, Agarwal & Greenbaum, 2021
• Hemidactylus granchii Lanza, 1978
• Hemidactylus graniticolus Agarwal, Giri & Bauer, 2011
• Hemidactylus granosus Heyden, 1827
• Hemidactylus granti Boulenger, 1899
• Hemidactylus greeffii Bocage, 1886
• Hemidactylus gujaratensis Giri, Bauer, Vyas & Patil, 2009
• Hemidactylus hajarensis Carranza & Arnold, 2012
• Hemidactylus hannahsabinae Ceríaco, Agarwal, Marques & Bauer, 2020
• Hemidactylus hegdei Pal & Mirza, 2022^[13]
• Hemidactylus hemchandrai Dandge & Tiple, 2015
• Hemidactylus homoeolepis Blanford, 1881
• Hemidactylus hunae Deraniyagala, 1937
• Hemidactylus imbricatus (Bauer, Giri, Greenbaum, Jackman, Dharne & Shouche, 2008)
• Hemidactylus inexpectatus Carranza & Arnold, 2012
• Hemidactylus inintellectus Sindaco, Ziliani, Razzetti, Pupin & Grieco, 2009
• Hemidactylus isolepis Boulenger, 1895
• Hemidactylus ituriensis Schmidt, 1919
• Hemidactylus jubensis Boulenger, 1895
• Hemidactylus jumailiae Busais & Joger, 2011
• Hemidactylus kamdemtohami Bauer & Pauwels, 2002
• 坎盖雷蜥虎 Hemidactylus kangerensis Mirza, Bhosale & Patil, 2017^[14]
• Hemidactylus karenorum (Theobald, 1868)
• Hemidactylus kimbulae Amarasinghe, Karunarathna, Campbell, Madawala & De Silva, 2021
• Hemidactylus klauberi Scortecci, 1948
• Hemidactylus kolliensis Agarwal, Bauer, Giri & Khandekar, 2019^[8]
• Hemidactylus kundaensis Chirio & Trape, 2012
• Hemidactylus kurdicus Safaei-Mahroo, Ghaffari, Ghafoor & Amini, 2017^[15]
• Hemidactylus kushmorensis Murray, 1884
• Hemidactylus kyaboboensis Wagner, Leaché & Fujita, 2014
• Hemidactylus laevis Boulenger, 1901^[16]
• Hemidactylus lamaensis Ullenbruch, Grell & Böhme, 2010
• Hemidactylus lankae Deraniyagala, 1953
• Hemidactylus lanzai Šmíd, Mazuch, Nováková, Modry, Malonza, Abdirahman-Elmi, Carranza & Moravec, 2020
• Hemidactylus laticaudatus Andersson, 1910
• Hemidactylus lavadeserticus Moravec & Böhme, 1997
• Hemidactylus lemurinus Arnold, 1980
• Hemidactylus leschenaultii Duméril & Bibron, 1836
• Hemidactylus longicephalus Bocage, 1873
• Hemidactylus lopezjuradoi Arnold, Vasconcelos, Harris, Mateo & Carranza, 2008
• Hemidactylus luqueorum Carranza & Arnold, 2012
• Hemidactylus mabouia (Moreau De Jonnès, 1818)
• Hemidactylus macropholis Boulenger, 1896
• Hemidactylus maculatus Duméril & Bibron, 1836
• Hemidactylus mahonyi Adhikari, Achyuthan, Kumar, Khot, Shreeram & Ganesh, 2022
• Hemidactylus makolowodei Bauer, Lebreton, Chirio, Ineich & Talla Kouete, 2006
• Hemidactylus malcolmsmithi (Constable, 1949)^[17]
• Hemidactylus mandebensis Šmíd, Moravec, Kratochv Il, Nasher, Mazuch, Gvoždík & Carranza, 2015
• Hemidactylus masirahensis Carranza & Arnold, 2012
• Hemidactylus matschiei (Tornier, 1901)
• Hemidactylus megalops Parker, 1932
• Hemidactylus mercatorius Gray, 1842
• Hemidactylus mindiae Baha El Din, 2005
• Hemidactylus minutus Vasconcelos & Carranza, 2014^[18]
• Hemidactylus modestus (Günther, 1894)
• Hemidactylus montanus Busais & Joger, 2011
• Hemidactylus mrimaensis Malonza & Bauer, 2014
• Hemidactylus multisulcatus Sayyed, Kirubakaran, Khot, Adhikari, Sayyed, Sayyed, Purkayastha, Deshpande & Sulakhe, 2023^[19]
• Hemidactylus muriceus Peters, 1870
• Hemidactylus murrayi Gleadow, 1887
• 纽顿蜥虎 Hemidactylus newtoni Ferreira, 1897
• Hemidactylus nicolauensis Vasconcelos, Köhler, Geniez & Crochet, 2020
• Hemidactylus nzingae Ceríaco, Agarwal, Marques & Bauer, 2020
• Hemidactylus ophiolepis Boulenger, 1903
• Hemidactylus ophiolepoides Lanza, 1978
• Hemidactylus oxyrhinus Boulenger, 1899
• Hemidactylus paaragowli Srikanthan, Swamy, Mohan & Pal, 2018^[20]
• Hemidactylus paivae Ceríaco, Agarwal, Marques & Bauer Bauer, 2020
• Hemidactylus pakkamalaiensis Narayanan, Christopher, Raman, Mukherjee, Vimalraj & Deepak^[21]
• Hemidactylus palaichthus Kluge, 1969
• Hemidactylus parvimaculatus Deraniyagala, 1953
• Hemidactylus pauciporosus Lanza, 1978
• Hemidactylus paucituberculatus Carranza & Arnold, 2012
• Hemidactylus persicus Anderson, 1872^[12]
• Hemidactylus pfindaensis Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Pinto, 2021
• Hemidactylus pieresii Kelaart, 1852
• Hemidactylus platycephalus Peters, 1854
• 蝎虎 Hemidactylus platyurus (Schneider, 1797)，蝎尾蜥虎
• Hemidactylus prashadi Smith, 1935
• Hemidactylus principensis Miller, Sellas & Drewes, 2012
• Hemidactylus pseudomuriceus Henle & Böhme, 2003
• Hemidactylus pseudoromeshkanicus Torki, 2019
• Hemidactylus puccionii Calabresi, 1927
• Hemidactylus pumilio Boulenger, 1899
• Hemidactylus quartziticolus Akshay Khandekar, Tejas Thackeray, Rameshwaran Mariappan, Satpal Gangalmale, Vivek Waghe, Swapnil Pawar and Ishan Agarwal. 2023. ^[22][23]
• Hemidactylus raya Kumar, Srinivasulu & Srinivasulu, 2022^[5]
• Hemidactylus reticulatus Beddome, 1870
• 理查德森蜥虎 Hemidactylus richardsonii (Gray, 1845)
• Hemidactylus rishivalleyensis Agarwal, Thackeray & Khandekar, 2020
• 强壮蜥虎 Hemidactylus robustus Heyden, 1827^[12]
• Hemidactylus romeshkanicus Torki, 2011
• Hemidactylus ruspolii Boulenger, 1896
• Hemidactylus saba Busais & Joger, 2011
• Hemidactylus sahgali Mirza, Gowande, Patil, Ambekar & Patel, 2018^[24]
• Hemidactylus sankariensis Agarwal, Bauer, Giri & Khandekar, 2019^[8]
• Hemidactylus sassanidianus Torki, 2019
• Hemidactylus sataraensis Giri & Bauer, 2008
• Hemidactylus saxicolus Kumar, Srinivasulu & Srinivasulu, 2022
• Hemidactylus scabriceps (Annandale, 1906)
• Hemidactylus shihraensis Busais & Joger, 2011
• Hemidactylus sinaitus Boulenger, 1885
• Hemidactylus sirumalaiensis Khandekar, Thackeray, Pawar & Agarwal, 2020
• Hemidactylus siva Srinivasulu, Srinivasulu & Kumar, 2018
• 史密斯蜥虎 Hemidactylus smithi Boulenger, 1895
• Hemidactylus somalicus Parker, 1932
• Hemidactylus squamulatus Tornier, 1896
• Hemidactylus srikanthani Adhikari, Achyuthan, Kumar, Khot, Shreeram & Ganesh, 2022
• 台湾蜥虎 Hemidactylus stejnegeri Ota & Hikida, 1989，史氏蜥虎
• Hemidactylus sushilduttai Giri, Bauer, Mohapatra, Srinvasulu & Agarwal, 2017^[25]
• Hemidactylus tamhiniensis Khandekar, Thackeray & Agarwal, 2021
• Hemidactylus tanganicus Loveridge, 1929
• Hemidactylus tasmani Hewitt, 1932
• 泰勒蜥虎 Hemidactylus taylori Parker, 1932
• Hemidactylus tenkatei Lidth De Jeude, 1895
• Hemidactylus thayene Zug & Mcmahan, 2007
• Hemidactylus treutleri Mahony, 2009
• Hemidactylus triedrus (Daudin, 1802)^[24]
• Hemidactylus tropidolepis Mocquard, 1888
• Hemidactylus turcicus (Linnaeus, 1758)
• Hemidactylus ulii Šmíd, Moravec, Kratochvíl, Gvoždík, Nasher, Busais, Wilms, Shobrak & Carranza, 2013
• Hemidactylus vanam Chaitanya, Lajmi & Giri, 2018^[26]
• Hemidactylus varadgirii Chaitanya, Agarwal, Lajmi & Khandekar, 2019
• Hemidactylus vernayi Ceríaco, Agarwal, Marques & Bauer, 2020
• 越南蜥虎 Hemidactylus vietnamensis Darevsky, Kupriyanova & Roshchin, 1984
• Hemidactylus vijayraghavani Mirza, 2018
• Hemidactylus whitakeri Mirza, Gowande, Patil, Ambekar & Patel, 2018^[24]
• Hemidactylus xericolus Lajmi, Giri, Singh & Agarwal, 2020^[11]
• Hemidactylus yajurvedi Murthy, Bauer, Lajmi, Agarwal & Giri, 2015^[27]
• Hemidactylus yerburii Anderson, 1895

参考文献

1. Hemidactylus Goldfuss, 1820. GBIF. [2021-07-03]. 
2. 赵尔宓，赵肯堂，周开亚等. 中国科学院中国动物志编辑委员会 , 编. . 北京: 科学出版社. 1999: 54-55 [2021-07-03]. ISBN 7-03-006750-9. （原始内容存档于2021-06-26）. 
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4. Zeeshan A Mirza, Rajesh V Sanap. 2014. A New Cryptic Species of Gecko of the genus Hemidactylus Oken, 1817 (Reptilia: Gekkonidae) from Southern India. TAPROBANICA. 6(1); 12-20. DOI: http://dx.doi.org/10.4038/tapro.v6i1.7056
5. Gandla Chethan Kumar, Aditya Srinivasulu and Chelmala Srinivasulu. 2022. Redescription of Hemidactylus giganteus Stoliczka, 1871 with the Description of Three New Allied Species (Squamata: Gekkonidae: Hemidactylus Goldfuss, 1820) from peninsular India. Zootaxa. 5115(3); 301-341. DOI: 10.11646/zootaxa.5115.3.1
6. Jiří Šmíd, Marek Uvizl, Mohammed Shobrak, Salem Busais, Al Faqih Ali Salim, Raed Hamoud M. AlGethami, Abdulaziz Raqi AlGethami, Abdulkarim Saleh K. Alanazi, Saad Dasman Alsubaie, Michail Rovatsos, Lucie Nováková, Tomáš Mazuch and Salvador Carranza. 2022. Diversification of Hemidactylus Geckos (Squamata: Gekkonidae) in coastal Plains and Islands of southwestern Arabia with Descriptions and Complete Mitochondrial Genomes of Two Endemic Species to Saudi Arabia. Organisms Diversity & Evolution. DOI: 10.1007/s13127-022-00572-w, http://Researchgate.net/publication/362741282_Diversification_Hemidactylus_Arabia
7. Jiří Šmíd, Mohammed Shobrak, Thomas M. Wilms, Ulrich Joger and Salvador Carranza. 2016. Endemic Diversification in the Mountains: Genetic, Morphological, and Geographical Differentiation of the Hemidactylus Geckos in southwestern Arabia. Organisms Diversity & Evolution. DOI: 10.1007/s13127-016-0293-3
8. Ishan Agarwal, Aaron B. Bauer, Varad B. Giri and Akshay Khandekar. 2019. An Expanded ND2 Phylogeny of the brookii and prashadi Groups with the Description of Three New Indian Hemidactylus Oken (Squamata: Gekkonidae). Zootaxa. 4619(3); 431–458. DOI: 10.11646/zootaxa.4619.3.2
9. Zeeshan A. Mirza and David Raju. 2017. A New Rupicolous Species of Gecko of the Genus Hemidactylus Oken, 1817 from the Satpura Hills, Central India. Amphibian & Reptile Conservation. 11(1) [General Section]: 51–71 (e137).
10. Philipp Wagner, Adam D. Leaché & Matthew K. Fujita. 2014. Description of Four New West African Forest Geckos of the Hemidactylus fasciatus Gray, 1842 complex, revealed by coalescent species delimitation. Bonn zoological Bulletin. 63 (1): 1–14
11. Aparna Lajmi, Varad B. Giri, Taneraw Singh and Ishan Agarwal. 2020. Two New Species of Yellow-tailed Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Rocky Outcrops on the Telangana Plateau, India. Zootaxa. 4895(4); 483–504. DOI: 10.11646/zootaxa.4895.4.2
12. Dan Cogălniceanu, Aurora M Castilla, Aitor Valdeón, Alberto Gosá, Noora Al-Jaidah, Ali Alkuwary, Essam O. H. Saifelnasr, Paloma Mas-Peinado, Renee Richer and Ahmad Amer Mohd Al-Hemaidi. 2014. A Preliminary Report on the Distribution of Lizards in Qatar. ZooKeys. 373: 67–91. doi: dx.doi.org/10.3897/zookeys.373.5994
13. Saunak Pal and Zeeshan A. Mirza. 2022. A New Species of Large-Bodied Gecko of The Genus Hemidactylus Goldfuss, 1820 from Southern Western Ghats of Tamil Nadu, India. Journal of Bombay Natural History Society. 119; DOI: 10.17087/jbnhs/2022/v119/167364
14. Zeeshan A. Mirza, Harshal Bhosale and Rishikesh Patil. 2017. A New Large Species of Gecko of the Genus Hemidactylus Oken, 1817 (Reptilia: Sauria: Gekkonidae) from the Eastern Ghats, India. Comptes Rendus Biologies. In Press. DOI: 10.1016/j.crvi.2017.09.003
15. Barbod Safaei-Mahroo, Hanyeh Ghaffari, Aram Ghafoor and Saywan Amini. 2017. A New Species of Hemidactylus (Squamata: Gekkota: Gekkonidae) from Qara Dagh Mountains, Kurdistan Region, with A Key to the Genus in Iraq. Zootaxa. 4363(3); 377–392. DOI: 10.11646/zootaxa.4363.3.4
16. Tomas Mazuch, Jiri Smid and Aaron M. Bauer. 2016. Rediscovery and A New Record of Hemidactylus laevis (Reptilia: Gekkonidae) from Somaliland, with Notes on and Resurrection of Hemidactylus fragilis.  Zootaxa. 4117(4):529. DOI: 10.11646/zootaxa.4117.4.5
17. Ishan Agarwal, Varad B. Giri and Aaron M. Bauer. 2018. On the Status of Cyrtodactylus malcolmsmithi (Constable, 1949). Breviora. 557; 1-11. DOI: 10.3099/MCZ41.1
18. Raquel Vasconcelos and Salvador Carranza. 2014. Systematics and Biogeography of Hemidactylus homoeolepis Blanford, 1881 (Squamata: Gekkonidae), with the Description of A New Species from Arabia. Zootaxa. 3835(4): 501–527. DOI: 10.11646/zootaxa.3835.4.4
19. Amit Sayyed, Samson Kirubakaran, Rahul Khot, Omkar Adhikari, Ayaan Sayyed, Masum Sayyed, Jayaditya Purkayastha, Shubhankar Deshpande and Shauri Sulakhe. 2023. A New Species of the Genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Tamil Nadu, India. Asian Journal of Conservation Biology. 12(1); 100–110. DOI: doi.org/10.53562/ajcb.82582
20. Achyuthan N. Srikanthan, Priyanka Swamy, Ashwini V. Mohan and Saunak Pal. 2018. A Distinct New Species of Riparian Rock-dwelling Gecko (Genus: Hemidactylus) from the southern Western Ghats. Zootaxa. 4434(1); 141–157. DOI: 10.11646/zootaxa.4434.1.9
21. Surya Narayanan, Peter Christopher, Kothandapani Raman, Nilanjan Mukherjee, Ponmudi Prabhu, Maniezhilan Lenin, Sivangnanaboopathidoss Vimalraj and V. Deepak. 2023. A New Species of Rock-dwelling Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the southern Eastern Ghats, India. Vertebrate Zoology. 73: 499-512. DOI: 10.3897/vz.73.e104494
22. A Remarkable New Species of Gecko (Squamata: Gekkonidae: Hemidactylus) from scrublands at the southern Tip of India. Vertebrate Zoology 73: 433-450. DOI: 10.3897/vz.73.e101871
23. 潜伏在印度寺庙墙壁上的刺状生物原来是一个新物种. 通樱科技网. [2023-05-22]. （原始内容于2023-05-22）. 
24. Zeeshan A. Mirza, Gaurang G. Gowande, Rishikesh Patil, Mayuresh Ambekar and Harshil Patel. 2018. First Appearance Deceives Many: Disentangling the Hemidactylus triedrus species complex Using An Integrated Approach. PeerJ. 6:e5341 DOI: 10.7717/peerj.5341
25. Varad B. Giri, Aaron M. Bauer, Pratyush P. Mohapatra, Chelmala Srinivasulu and Ishan Agarwal. 2017. A New Species of Large-bodied, Tuberculate Hemidactylus Oken (Squamata: Gekkonidae) from the Eastern Ghats, India. Zootaxa. 4347(2); 331–345. DOI: 10.11646/zootaxa.4347.2.8
26. R. Chaitanya, Aparna Lajmi and Varad B. Giri. 2018. A New Cryptic, Rupicolous Species of Hemidactylus Oken, 1817 (Squamata: Gekkonidae) from Meghamalai, Tamil Nadu, India. Zootaxa. 4374(1); 49–70. DOI: 10.11646/zootaxa.4374.1.3
27. B.H.C.K. Murthy, A. M. Bauer, Aparna Lajmi, Ishan Agarwal and Varad Giri. 2015. A New Rock Dwelling Hemidactylus (Squamata: Gekkonidae) from Chhattisgarh, India. Zootaxa. 4021(2): 334–350. DOI: 10.11646/zootaxa.4021.2.5